Gaster (1924, 1927, 1929, 1932, 1937, 1941, 1944) - Actinocamax plenus Subzone

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Biostratigraphical correlation: Equates with the greater part of the M. geslinianum Zone of current usage.

Lithostratigraphical correlation: Equates with the Plenus Marls Member, at the base of the Holywell Nodular Chalk Formation in southern England.

Age: Late Cenomanian