Gaster (1924, 1927, 1929, 1932, 1937, 1941, 1944) - Actinocamax quadratus Zone

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Biostratigraphical correlation: Equates with the middle and higher parts of the G. quadrata Zone (A. cretaceus Subzone & post Applinocrinus beds) of current usage.

Lithostratigraphical correlation: Equates with the middle and higher parts of the Culver Chalk Formation and basal Portsdown Chalk Formation of southern England.

Subzones: Saccocoma cretacea

Age: Campanian