Gaster (1924, 1927, 1929, 1932, 1937, 1941, 1944) - Echinocorys depressula Subzone

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Biostratigraphical correlation: Equates with the upper U. anglicus Zone and E depressula Subzone of current usage (Young & Lake, 1988).

Lithostratigraphical correlation: Equates with the lower to middle Newhaven Chalk Formation of southern England.

Age: Campanian