Gaster (1924, 1927, 1929, 1932, 1937, 1941, 1944) - Marsupites testudinarius Zone

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Biostratigraphical correlation: Equates with the U. socialis, M. testudinarius and U. anglicus Zone (pars) of current usage (Young & Lake, 1988).

Lithostratigraphical correlation: Equates with the lower part of the Newhaven Chalk Formation of southern England.

Age: Late Santonian & Early Campanian