Griffith & Brydone (1911); Brydone (1912) - Holaster planus Zone

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Biostratigraphical correlation: Inferred to be equivalent to Jukes-Browne & Hill's (1904) concept of this zone, and therefore probably equivalent to an interval from the base of the S. plana Zone to within the top of the S. plana Zone of current usage.

Lithostratigraphical correlation: Inferred to equate with the lower to middle part of the Lewes Nodular Chalk of southern England.

Age: Late Turonian