Griffith & Brydone (1911); Brydone (1912) - Marsupites Zone

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Biostratigraphical correlation: Equates with the M. testudinarius Zone and part of the U. anglicus Zone of current usage (Young & Lake, 1988).

Lithostratigraphical correlation: Equates with an interval in the lower part of the Newhaven Chalk of southern England.

Age: Santonian & basal Campanian