BGS Zone 19 (=upper UKB15 of Hart et al., 1989)
Base: Inception of Bolivinoides culverensis.
Top: Inception of Gavelinella usakensis.
Biomarkers:
- Inception of very rare Globotruncana arca at the base of the foraminiferal zone.
- Extinction of Globigerinelloides rowei in the basal O. pilula macrofaunal zone.
- Extinction of Stensioeina exsculpta exsculpta in the earliest O. pilula macrofaunal zone.
- Stensioeina pommerana inception in the mid O. pilula macrofaunal zone (transitional specimens closely resembling this species occur in the late Santonian and early Campanian).
- Eouvigerina gracilis and Reussella kelleri extinction immediately below the Arundel Sponge Bed and lateral equivalents.
Macrofaunal correlation: Lowest O. pilula Zone to lowermost G. quadrata Zone.
Age: Campanian