Griffith & Brydone (1911); Brydone (1912) - E. depressula Subzone
Biostratigraphical correlation: Equates to an interval from within the U. anglicus Zone to a little above the Subzone of abundant O. pilula of current usage (Young & Lake, 1988).
Lithostratigraphical correlation: Equates with the middle part of the Newhaven Chalk Formation of southern England.
Age: Campanian