Griffith & Brydone (1911); Brydone (1912) - E. depressula Subzone

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Biostratigraphical correlation: Equates to an interval from within the U. anglicus Zone to a little above the Subzone of abundant O. pilula of current usage (Young & Lake, 1988).

Lithostratigraphical correlation: Equates with the middle part of the Newhaven Chalk Formation of southern England.

Age: Campanian

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