Griffith & Brydone (1911); Brydone (1912) - Marsupites Zone
Biostratigraphical correlation: Equates with the M. testudinarius Zone and part of the U. anglicus Zone of current usage (Young & Lake, 1988).
Lithostratigraphical correlation: Equates with an interval in the lower part of the Newhaven Chalk of southern England.
Age: Santonian & basal Campanian