Griffith & Brydone (1911); Brydone (1912) - abundant O. pilula Subzone

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Biostratigraphical correlation: Equates with an interval from within the E. depressula Subzone to just below the top of the Subzone of abundant O. pilula of current usage (Young & Lake, 1988).

Lithostratigraphical correlation: Equates with the upper part of the Newhaven Chalk and, locally, the basal part of the Culver Chalk Formation of southern England.

Age: Campanian